Caporegime
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Bees are flying insects closely related to wasps and ants, and are known for their role in pollination, and in the case of the best-known bee species, the European honey bee, for producing honey and beeswax. Bees are a monophyletic lineage within the superfamily Apoidea, presently considered as a clade Anthophila. There are nearly 20,000 known species of bees in seven to nine recognized families,[1] though many are undescribed and the actual number is probably higher. They are found on every continent except Antarctica, in every habitat on the planet that contains insect-pollinated flowering plants.
Some species including honey bees, bumblebees, and stingless bees live socially in colonies. Bees are adapted for feeding on nectar and pollen, the former primarily as an energy source and the latter primarily for protein and other nutrients. Most pollen is used as food for larvae. Bee pollination is important both ecologically and commercially; the decline in wild bees has increased the value of pollination by commercially managed hives of honey bees.
Bees range in size from tiny stingless bee species whose workers are less than 2 millimetres (0.08 in) long, to Megachile pluto, the largest species of leafcutter bee, whose females can attain a length of 39 millimetres (1.54 in). The most common bees in the Northern Hemisphere are the Halictidae, or sweat bees, but they are small and often mistaken for wasps or flies. Vertebrate predators of bees include birds such as bee-eaters; insect predators include beewolves and dragonflies.
Human beekeeping or apiculture has been practised for millennia, since at least the times of Ancient Egypt and Ancient Greece. Apart from honey and pollination, honey bees produce beeswax, royal jelly and propolis. Bees have appeared in mythology and folklore, again since ancient times, and they feature in works of literature as varied as Beatrix Potter's The Tale of Mrs Tittlemouse, W. B. Yeats's poem The Lake Isle of Innisfree, and Laline Paull's The Bees. Bee larvae are included in the Javanese dish botok tawon, where they are eaten steamed with shredded coconut.
Did you know that:
The ancestors of bees were wasps in the family Crabronidae, which were predators of other insects. The switch from insect prey to pollen may have resulted from the consumption of prey insects which were flower visitors and were partially covered with pollen when they were fed to the wasp larvae. This same evolutionary scenario may have occurred within the vespoid wasps, where the pollen wasps evolved from predatory ancestors. Until recently, the oldest non-compression bee fossil had been found in New Jersey amber, Cretotrigona prisca of Cretaceous age, a corbiculate bee.[2] A bee fossil from the early Cretaceous (~100 mya), Melittosphex burmensis, is considered "an extinct lineage of pollen-collecting Apoidea sister to the modern bees".[3] Derived features of its morphology (apomorphies) place it clearly within the bees, but it retains two unmodified ancestral traits (plesiomorphies) of the legs (two mid-tibial spurs, and a slender hind basitarsus), showing its transitional status.[3] By the Eocene (~45 mya) there was already considerable diversity among eusocial bee lineages.[4][a]
The highly eusocial corbiculate Apidae appeared roughly 87 Mya, and the Allodapini (within the Apidae) around 53 Mya.[7] The Colletidae appear as fossils only from the late Oligocene (~25 Mya) to early Miocene.[8] The Melittidae are known from Palaeomacropis eocenicus in the Early Eocene.[9] The Megachilidae are known from trace fossils (characteristic leaf cuttings) from the Middle Eocene.[10] The Andrenidae are known from the Eocene-Oligocene boundary, around 34 Mya, of the Florissant shale.[11] The Halictidae first appear in the Early Eocene[12] with species [13][14] found in amber. The Stenotritidae are known from fossil brood cells of Pleistocene age.[15]
So short story - bees are awesome.
It is usually easy to recognise that a particular insect is a bee. They differ from closely related groups such as wasps by having branched or plume-like setae (bristles), combs on the forelimbs for cleaning their antennae, small anatomical differences in the limb structure and the venation of the hind wings, and in females, by having the seventh dorsal abdominal plate divided into two half-plates.[22]
Behaviourally, one of the most obvious characteristics of bees is that they collect pollen to provide provisions for their young, and have the necessary adaptations to do this. However, certain wasp species such as pollen wasps have similar behaviours, and a few species of bee scavenge from carcases to feed their offspring.[22] The world's largest species of bee is thought to be the Indonesian resin bee Megachile pluto, whose females can attain a length of 39 millimetres (1.54 in).[23] The smallest species may be dwarf stingless bees in the tribe Meliponini whose workers are less than 2 millimetres (0.08 in) in length.[24]
Head-on view of a carpenter bee, showing antennae, three ocelli, compound eyes, sensory bristles and mouthparts
A bee has a pair of large compound eyes which cover much of the surface of the head. Between and above these are three small simple eyes (ocelli) which provide information for the bee on light intensity. The antennae usually have thirteen segments in males and twelve in females and are geniculate, having an elbow joint part way along. They house large numbers of sense organs that can detect touch (mechanoreceptors), smell and taste, and small, hairlike mechanoreceptors that can detect air movement so as to "hear" sounds. The mouthparts are adapted for both chewing and sucking by having both a pair of mandibles and a long proboscis for sucking up nectar.[25]
The thorax has three segments, each with a pair of robust legs, and a pair of membranous wings on the hind two segments. The front legs of corbiculate bees bear combs for cleaning the antennae, and in many species the hind legs bear pollen baskets, flattened sections with incurving hairs to secure the collected pollen. The wings are synchronised in flight and the somewhat smaller hind wings connect to the forewings by a row of hooks along their margin which connect to a groove in the forewing. The abdomen has nine segments, the hindermost three being modified into the sting.[25]
Predators, parasites and pathogens
The bee-eater, Merops apiaster, specialises in feeding on bees; here a male catches a nuptial gift for his mate.
Further information: diseases of the honey bee
Vertebrate predators of bees include bee-eaters, shrikes and flycatchers, which make short sallies to catch insects in flight.[63] Swifts and swallows[63] fly almost continually, catching insects as they go. The honey buzzard attacks bees' nests and eats the larvae.[64] The greater honeyguide interacts with humans by guiding them to the nests of wild bees. The humans break open the nests and take the honey and the bird feeds on the larvae and the wax.[65] Among mammals, predators such as the badger dig up bumblebee nests and eat both the larvae and any stored food.[66]
The beewolf Philanthus triangulum paralysing a bee with its sting
Specialist ambush predators of visitors to flowers include crab spiders, which wait on flowering plants for pollinating insects; predatory bugs, and praying mantises,[63] some of which (the flower mantises) wait motionless, aggressive mimics camouflaged as flowers.[67] Beewolves are large wasps that habitually attack bees;[63] the ethologist Niko Tinbergen estimated that a single colony of the beewolf Philanthus triangulum might kill several thousand honeybees in a day: all the prey he observed were honeybees.[68] Other predatory insects that sometimes catch bees include robber flies and dragonflies.[63]
Honey bees are affected by parasites including acarine and Varroa mites.[69]
Tell me your bee thoughts please
Some species including honey bees, bumblebees, and stingless bees live socially in colonies. Bees are adapted for feeding on nectar and pollen, the former primarily as an energy source and the latter primarily for protein and other nutrients. Most pollen is used as food for larvae. Bee pollination is important both ecologically and commercially; the decline in wild bees has increased the value of pollination by commercially managed hives of honey bees.
Bees range in size from tiny stingless bee species whose workers are less than 2 millimetres (0.08 in) long, to Megachile pluto, the largest species of leafcutter bee, whose females can attain a length of 39 millimetres (1.54 in). The most common bees in the Northern Hemisphere are the Halictidae, or sweat bees, but they are small and often mistaken for wasps or flies. Vertebrate predators of bees include birds such as bee-eaters; insect predators include beewolves and dragonflies.
Human beekeeping or apiculture has been practised for millennia, since at least the times of Ancient Egypt and Ancient Greece. Apart from honey and pollination, honey bees produce beeswax, royal jelly and propolis. Bees have appeared in mythology and folklore, again since ancient times, and they feature in works of literature as varied as Beatrix Potter's The Tale of Mrs Tittlemouse, W. B. Yeats's poem The Lake Isle of Innisfree, and Laline Paull's The Bees. Bee larvae are included in the Javanese dish botok tawon, where they are eaten steamed with shredded coconut.
Did you know that:
The ancestors of bees were wasps in the family Crabronidae, which were predators of other insects. The switch from insect prey to pollen may have resulted from the consumption of prey insects which were flower visitors and were partially covered with pollen when they were fed to the wasp larvae. This same evolutionary scenario may have occurred within the vespoid wasps, where the pollen wasps evolved from predatory ancestors. Until recently, the oldest non-compression bee fossil had been found in New Jersey amber, Cretotrigona prisca of Cretaceous age, a corbiculate bee.[2] A bee fossil from the early Cretaceous (~100 mya), Melittosphex burmensis, is considered "an extinct lineage of pollen-collecting Apoidea sister to the modern bees".[3] Derived features of its morphology (apomorphies) place it clearly within the bees, but it retains two unmodified ancestral traits (plesiomorphies) of the legs (two mid-tibial spurs, and a slender hind basitarsus), showing its transitional status.[3] By the Eocene (~45 mya) there was already considerable diversity among eusocial bee lineages.[4][a]
The highly eusocial corbiculate Apidae appeared roughly 87 Mya, and the Allodapini (within the Apidae) around 53 Mya.[7] The Colletidae appear as fossils only from the late Oligocene (~25 Mya) to early Miocene.[8] The Melittidae are known from Palaeomacropis eocenicus in the Early Eocene.[9] The Megachilidae are known from trace fossils (characteristic leaf cuttings) from the Middle Eocene.[10] The Andrenidae are known from the Eocene-Oligocene boundary, around 34 Mya, of the Florissant shale.[11] The Halictidae first appear in the Early Eocene[12] with species [13][14] found in amber. The Stenotritidae are known from fossil brood cells of Pleistocene age.[15]
So short story - bees are awesome.
It is usually easy to recognise that a particular insect is a bee. They differ from closely related groups such as wasps by having branched or plume-like setae (bristles), combs on the forelimbs for cleaning their antennae, small anatomical differences in the limb structure and the venation of the hind wings, and in females, by having the seventh dorsal abdominal plate divided into two half-plates.[22]
Behaviourally, one of the most obvious characteristics of bees is that they collect pollen to provide provisions for their young, and have the necessary adaptations to do this. However, certain wasp species such as pollen wasps have similar behaviours, and a few species of bee scavenge from carcases to feed their offspring.[22] The world's largest species of bee is thought to be the Indonesian resin bee Megachile pluto, whose females can attain a length of 39 millimetres (1.54 in).[23] The smallest species may be dwarf stingless bees in the tribe Meliponini whose workers are less than 2 millimetres (0.08 in) in length.[24]
Head-on view of a carpenter bee, showing antennae, three ocelli, compound eyes, sensory bristles and mouthparts
A bee has a pair of large compound eyes which cover much of the surface of the head. Between and above these are three small simple eyes (ocelli) which provide information for the bee on light intensity. The antennae usually have thirteen segments in males and twelve in females and are geniculate, having an elbow joint part way along. They house large numbers of sense organs that can detect touch (mechanoreceptors), smell and taste, and small, hairlike mechanoreceptors that can detect air movement so as to "hear" sounds. The mouthparts are adapted for both chewing and sucking by having both a pair of mandibles and a long proboscis for sucking up nectar.[25]
The thorax has three segments, each with a pair of robust legs, and a pair of membranous wings on the hind two segments. The front legs of corbiculate bees bear combs for cleaning the antennae, and in many species the hind legs bear pollen baskets, flattened sections with incurving hairs to secure the collected pollen. The wings are synchronised in flight and the somewhat smaller hind wings connect to the forewings by a row of hooks along their margin which connect to a groove in the forewing. The abdomen has nine segments, the hindermost three being modified into the sting.[25]
Predators, parasites and pathogens
The bee-eater, Merops apiaster, specialises in feeding on bees; here a male catches a nuptial gift for his mate.
Further information: diseases of the honey bee
Vertebrate predators of bees include bee-eaters, shrikes and flycatchers, which make short sallies to catch insects in flight.[63] Swifts and swallows[63] fly almost continually, catching insects as they go. The honey buzzard attacks bees' nests and eats the larvae.[64] The greater honeyguide interacts with humans by guiding them to the nests of wild bees. The humans break open the nests and take the honey and the bird feeds on the larvae and the wax.[65] Among mammals, predators such as the badger dig up bumblebee nests and eat both the larvae and any stored food.[66]
The beewolf Philanthus triangulum paralysing a bee with its sting
Specialist ambush predators of visitors to flowers include crab spiders, which wait on flowering plants for pollinating insects; predatory bugs, and praying mantises,[63] some of which (the flower mantises) wait motionless, aggressive mimics camouflaged as flowers.[67] Beewolves are large wasps that habitually attack bees;[63] the ethologist Niko Tinbergen estimated that a single colony of the beewolf Philanthus triangulum might kill several thousand honeybees in a day: all the prey he observed were honeybees.[68] Other predatory insects that sometimes catch bees include robber flies and dragonflies.[63]
Honey bees are affected by parasites including acarine and Varroa mites.[69]
Tell me your bee thoughts please
