1x4gb or 2x2gb ?

4x1 will work the same, but since the memory controller has more sticks to work with, it will be more stressed and may require more voltages.
 
You'd be able to take advantage of dual channel (still), the dimms will pair up, however it'll likely require more voltage to drive 4 slots than it will 2. Not an issue if you're not trying to overclock to limits usually, but it is less efficient and obviously cramps your capability to upgrade further.

Edit: double sniped..!
 
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