There appears to be a difference in how the PCIEX4 slot shares bandwidth. On the cheaper board, Z77X-UD3H, this shares bandwidth with the PCIe x1 slots. When a PCIe x4 card is installed in PCIEX4 all PCIe x1 slots become unavailable. In contrast the more expensive Z77X-UD5H, the PCIEX4 slot shares bandwidth with the other two PCI Express x16 slots so whilst those are reduced in bandwidth as you'd expect, it doesn't disable the PCIe x1 slots.