Evolution in Action:
This section is simply to expand on knowledge that one may know on this subject to gain a greater, more educated understanding. This could go on for a very long time, but I shall keep it relatively brief.
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The 4 Postulates of Evolution by Natural Selection:
Natural selection produces descent with modification, i.e. evolution and the following postulations describe this method. All postulates are testable.
1) Individuals within species are variable (because of mutation creating new alleles and subsequent shuffling of alleles).
2) Some of the variations are passed on to offspring (genes are passed on to offspring intact and often independently of other genes).
3) In most generations, more offspring are produced than can survive.
4) Survival and reproduction are not random: individuals with the highest reproductive success are those with the most favourable variations (with alleles and allelic combinations that best adapt them to their environment)- they are ‘naturally selected’.
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Darwin’s Finches – Testing the postulates of natural selection:
Darwin’s finches are a collection of birds from the Galapagos Islands, a series of islands which have been largely undisturbed by human activity and show mass biological diversity. They evolved by allopatric speciation (more later) and adaptive radiation, probably evolved from common ancestor 1-5 Mya. Throughout the 70s and 80s, the Grant team observed and tagged individuals on the islet of Daphne Major, allowing each of these postulates to be tested and proved.
Postulate 1 - Individuals within species are variable:
The normal distribution of bill depth for Geospiza fortis:
The chart shows that there is clearly variation between individual bill depth. A drought in 1977-1978 caused a huge population crash, shown by the second chart. The drought caused a reduction in small seeds, leaving only seeds which birds with large bill depth could consume.
Postulate 2 - Some of the variations are passed on to offspring:
Regression analysis shows how much the change of a variable is due to one factor. The regression slope of the following chart measures 0.79 (R2 statistic), means that 79% of the variation in bill depth due to the additive effects of genes.
Postulate 3 - Are more organisms born than survive to reproduce?
The huge population crash in 1977 coincided with severe drought (Giving 85% mortality). Most finches died of starvation when seeds became scarce, however some population did surive
Postulate 4 - is survival and reproduction nonrandom??
Yes. There was a survival bias towards birds with deeper bills which could break open the hard seeds of Tribulus cistodes which were still relatively abundant. Only birds with deep bills can crack these seeds, so large bills selected for. This is shown in the chart below:
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The Three Types of Selection:
Directional selection: Fitness increases (or decreases) with trait magnitude.
Stabilising selection: Individuals with intermediate trait values have highest fitness.
Disruptive selection: Individuals with extreme trait values have greatest fitness. This may be important in some modes of speciation.
The three types of selection demonstrated below: