The PLX chip adds extra lanes to the bottom PCI_E 16X slot meaning it can run at upto 4X bandwidth without affecting the performance of say the USB controllers/SATA controllers or in the case of the PRO board,
http://www.asus.com/Motherboards/Intel_Socket_1155/P8Z68V_PRO/#specifications
The two top 16X slots still run at 8X/8X just like the Deluxe does.
The bottom slots performance has no barring on what you are wanting to do. Just dont use it or you may find the triple slot cooler of the ASUS directCUII card will block it if you get an identical card or use this bigger card in the second 16X slot while using a more normal dual slot thickness GTX580 in the top 16X slot.
http://www.asus.com/Motherboards/Intel_Socket_1155/P8Z68V_PRO/#specifications
*1: The PCIe x16_3 slot shares bandwidth with PCIe x1_1 slot, PCIe x1_2 slot, USB3_34 and ESATA12. The PCIe x16_3 runs at x1 mode by default for system resource optimization.(PCIe x1_2 will be disabled.)
The two top 16X slots still run at 8X/8X just like the Deluxe does.
The bottom slots performance has no barring on what you are wanting to do. Just dont use it or you may find the triple slot cooler of the ASUS directCUII card will block it if you get an identical card or use this bigger card in the second 16X slot while using a more normal dual slot thickness GTX580 in the top 16X slot.
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